The basic way you tune a cocker or sterling is the same
Changes in velocity are achived by swapping springs (hammer and valve), shimming in the case of older Stirlings, or using the velocity dialer (IVG) to change hammer spring tension. The main difference is the hammer spring is dialled from the rear on cockers, and from the front through the pump on later sterlings.
The inline regulator is used to set input pressure, and make a marker more consistent over chronograph. This should ideally be sweet spotted to match spring set. You do this by increasing input pressure until velocity starts to drop off, then back off input pressure to the point the highest fps reading was achieved.
Old markers were originally set up to run on CO2, and won't hit usable velocity on HP air without a secondary regulator. CO2 bottles run at around 650-700psi, compared to HP air at 850 psi. The higher pressure will hold the cup seal in the valve shut, fighting against spring tension and capping velocity